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A study where this was investigated in hepatocytes expressing CYP2E1 in both locations or exclusively in mitochondria showed a range of adverse effects in both types of cell (despite lower cellular CYP2E1 in the latter)

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If the stress persists, ATF4 can also promote the transcription of C/EBP homologous protein (CHOP), a pro-apoptotic protein, to induce cell apoptosis (57)