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As well as being a primary precursor of (-)-jasmonic acid, (+)-12-oxo-phytodienoic acid ( cis -OPDA) is reported to regulate growth processes and certain jasmonate-responsive genes in plants (although some doubts have been expressed), and it induces and fine-tunes defence and stress responses (see below), as does a C 20 analogue of OPDA detected in a liverwort